Giant Pill Millipede
Procyliosoma
Their Story
The giant pill millipede might look like a shiny dark cherry-sized ball—but when it unrolls at night, it reveals a multitude of legs quietly working the forest floor. Endemic to Aotearoa and surviving here in the Brook Waimārama Sanctuary’s pest-free native forest, its presence is a sign of ecological depth and recovery.
Conservation Status

Pill millipedes are considered not threatened with extinction, but rely on intact forest to flourish.
Food

Mostly nocturnal, giant pill millipedes unfurl at night to feed on decaying wood, fallen leaves and other detritus—playing a vital role in nutrient recycling.
These compact millipedes are distinguished by their ability to roll up into a near-perfect ball (called conglobation), hiding head and legs entirely beneath glossy plates. Each segment (tergite) overlaps the next, forming a protective shield. Colours vary from dark brown to black-shiny, with adult sizes comparable to a large marble or cherry.
Mostly nocturnal, giant pill millipedes unfurl at night to feed on decaying wood, fallen leaves and other detritus—playing a vital role in nutrient recycling. During the day they remain curled into protective balls beneath logs, bark or leaf-litter—a clever adaptation that shields them from predators and the elements.
Adults mature slowly and have relatively low reproductive output. Females lay eggs in moist forest litter or under logs. Hatchlings emerge as miniature adults and gradually grow through years of forest debris-rich habitat. Their longevity and slow life-history make intact forest and protected conditions crucial for their survival.
The genus Procyliosoma comprises three described species endemic to Aotearoa, with other species endemic to Australia and Tasmania. They live in moist, mature native forest rich in leaf-litter and rotting logs. Within the Brook Sanctuary’s beech-podocarp forest, the abundance of these millipedes reflects the Sanctuary’s old growth and organic substrate build-up.
Giant pill millipedes are not considered threatened with extinction, however these species have suffered declines due to the loss and degradation of mature forest. Their survival is enhanced inside pest-free sanctuaries like Brook Waimārama.
Pill millipedes are rarely spotted during the day, hiding beneath logs and litter. Come nightfall however, these strange invertebrates become conspicuous. They seem to be very common in the moister areas of the Sanctuary, becoming very large.
Millipedes are the modern relatives of an ancient group of arthropods known as Diplopods. These ‘double legged’ segmented invertebrates arrived on the scene more than 400 million years ago, tens of millions of years before vertebrates emerged onto land. The largest millipede(and terrestrial invertebrate) to ever live was Arthropleura of the Carboniferous period, which grew over 2 metres long
































